Alexandre Yersin identifies the plague bacterium
In 1894, during a plague outbreak in Hong Kong, Alexandre Yersin identified the bacterium that would later be named Yersinia pestis. This discovery transformed understanding of the Black Death by replacing older theories of poisoned air or divine punishment with a microbiological explanation. Although it occurred centuries after the medieval pandemic, it is a milestone in Black Death history because it provided the scientific basis for linking medieval plague to a specific pathogen. That breakthrough opened the way for modern epidemiology, laboratory diagnosis, and eventually historical DNA research into pandemic origins and spread.